Wet meadows of montane areas of the higher Hercynian ranges, the Jura and the Alpine piedmont rich in Trollius europaeus and Cirsium rivulare, replacing the cabbage thistle meadows of unit E3.412 at higher altitudes. Their cortège includes Cirsium oleraceum, Caltha palustris, Lychnis flos-cuculi, Myosotis palustris, Geum rivale, Festuca pratensis, Galium album, Ranunculus acris, Holcus lanatus and, sometimes, in particular in the Swiss Jura, Fritillaria meleagris.