More humid, or temporarily more humid, swards of the lowland sub-Atlantic mesophile hay meadows of Western Europe and Central Europe, dominated by Arrhenatherum elatius and Alopecurus pratensis, or by the latter alone, and with a species composition intermediate between that of mesophile and humid meadows (E3) with Cirsium oleraceum, Angelica sylvestris, Sanguisorba officinalis, Ranunculus repens, Myosotis palustris, Glechoma hederacea, Lychnis flos-cuculi, Ajuga reptans, Cardamine pratensis, Lysimachia nummularia, Geranium pratense, Campanula patula, Pastinaca sativa, Heracleum sphondylium, Anthriscus sylvestris. Towards the east, in more continental climates, the Alopecurus meadows communities increasingly take the character of humid riverine meadows; they have been included in E3 from the Pannonic region eastwards, in the range of the mesophile meadows of E2.15.