Sphagnum-rich Picea abies forests of the Alps, developed on more or less peaty, humid but not waterlogged substrates, with Listera cordata, Sphagnum acutifolium, Sphagnum quinquefarium, Sphagnum girgensohnii. Spruce forests developed in fens or swamps at the periphery of raised bogs or on waterlogged soils in forests are included in swamp forests of class G3, as unit G3.E5.