Regularly grazed mesotrophic pastures of Europe, fertilised and on well-drained soils, with Lolium perenne, Cynosurus cristatus, Poa spp., Festuca spp., Trifolium repens, Leontodon autumnalis, Bellis perennis, Ranunculus repens, Ranunculus acris, Cardamine pratensis, Deschampsia cespitosa; they are most characteristic of the nemoral and boreonemoral zones Europe, but extend to the Cordillera Central, the Apennines and the supra-Mediterranean zone of the Balkan peninsula and Greece.