Forests of Quercus robur, sometimes Quercus petraea, rich in Fraxinus excelsior, with Carpinus betulus, developed on more or less wet, meso-eutrophic soils, in regions of moderate Atlantic influence, from southern Champagne and Lorraine north to lower Rhenania, Westphalia and Hanover, characterized by the abundance of species of the ecological groups of Primula elatior, of Lamium galeobdolon, of Anemone nemorosa and by the absence of Hyacinthoides non-scripta.
Quercus robur-Fraxinus excelsior forests occupying damp colluvions at the bottom of slopes in valleys within sub-Atlantic forests, characterized by the presence of the group of Anemone ranunculoides, Corydalis solida, Gagea lutea and Lathraea squamaria or of Aconitum vulparia, transitional to ravine or alluvial forests.