Forests of Fraxinus excelsior, Acer pseudoplatanus, Acer platanoides, Ulmus glabra, Tilia platyphyllos, Fagus sylvatica, of calcareous block screes and rocky slopes, in shady, humid ravines of the hills, low mountains and plateaux associated with the Hercynian ranges, the Jura, the Paris Basin, the western, northern and locally eastern and southeastern pre-Alps, the Northern Carpathians, the Vertes, Bakony and Bükk hills of the Pannonic plain, characterized by the dominance in the understorey of Asplenium scolopendrium and the presence of Ribes uva-crispa, Asplenium trichomanes, Asplenium viride, Cystopteris fragilis, Polystichum aculeatum, Moehringia muscosa, Chrysosplenium alternifolium, Valeriana tripteris, Adenostyles alpina; accompanying subdominants are shared with other ravine forests, in particular, Mercurialis perennis, Lunaria rediviva, Lamiastrum galeobdolon ssp. montanum, Galium odoratum, Dryopteris filix-mas, Ctenidium molluscum.