Mats of Arctostaphylos uva-ursi or Arctostaphylos alpinus of the alpine, subalpine and locally, montane, belts of the Alps, the Pyrenees, the northern and central Apennines, the Dinarides, the Carpathians, the Balkan Range, the Rhodope Mountains (south to the Slavianka-Orvilos, the Menikion, the Pangeon, the Falakron and the Rhodopi), the Moeso-Macedonian mountains (including Athos), the Pelagonides (south to the Greek Macedonian border ranges Tzena, Pinovon and Kajmakchalan) and Olympus, in the Thessalian mountains, mostly on calcareous substrates.