Calcareous and calcschist screes of warm exposures in mountain ranges of the nemoral zone, including the Alps, Pyrenees and Caucasus, and of Mediterranean mountains, hills and lowlands and, locally, of warm, sunny middle European upland or lowland sites.
Supra-Mediterranean screes of Mediterranean southern France, common in the coastal ranges of the Marseille region (Allauch, Carpiagne, Puget, Marseilleveyre), rare on the Sainte-Baume, also represented in the Cévennes, with Arenaria provincialis (Gouffeia arenarioides), Ptychotis heterophylla, Linaria supina, Centranthus ruber, Centranthus lecoqii, Crucianella latifolia.