Medio-European and Atlantic forests, on neutral or near-neutral soils, with mild humus (mull), characterized by a strong representation of species belonging to the ecological groups of Anemone nemorosa, of Lamium galeobdolon, of Galium odoratum and Melica uniflora and, in mountains, various Dentaria, forming a richer and more abundant herb layer than in 41.11 and 41.12.
Medio-European collinar beech and beech-oak forests of the Hercynian arc and peripheral regions, the Jura, Lorraine, the Paris basin, Burgundy and a few localities of the North Sea - Baltic plain.