Medio-European and Atlantic forests, on neutral or near-neutral soils, with mild humus (mull), characterized by a strong representation of species belonging to the ecological groups of Anemone nemorosa, of Lamium galeobdolon, of Galium odoratum and Melica uniflora and, in mountains, various Dentaria, forming a richer and more abundant herb layer than in 41.11 and 41.12.